Subject: Caves: Into the Underworld
     The Nation
      Wed, April 1, 1998

Exploring the Underworld

      Blind fish. Hairy Mary's. Living
      rock. These are just some of
      the weird and wondrous things
      to be found in the still largely
      undiscovered caves of
      Thailand. James Fahn reports.

      SO you thought there weren't
      any places left in this
      ever-shrinking world to be
      explored.

      Richard Borowsky begs to
      differ. A flag-carrying member of
      the Explorers Club, he set off
      earlier this month with two
      colleagues to explore one of
      Thailand's last frontiers: Its
      caves.

      Plunging into Phra That Cave in
      Kanchanaburi's Erawan
      National Park is like entering a
      negative of the rich, forested
      world above. There are few
      colours here, and little life. Even
      the air can forsake you, as
      pockets of carbon dioxide
      occasionally make it difficult to
      breathe.

      But still there are plenty of
      marvels to be seen. Over
      millions of years, the interaction
      of water, air and limestone has
      created bizarre rock formations
      that lend the underworld an
      austere beauty.

      Huge flowstones of sensuously
      rounded rock spill down to the
      stream-bed. Tiny tendrils of
      limestone, creamy white like
      vanilla icing, drip down from the
      ceiling, each tipped with a bead
      of water, eventually to turn into
      large stalactites. Elsewhere the
      stone is shaped like curtains,
      and in one instance we come
      across large strips of striped
      rock which cavers dub ''bacon''.

      As you trudge, slip and slide
      through this muddy, watery
      world, it's easy to convince
      yourself that you are crawling
      through the Earth's innards.

      What life that does exist here
      seems out of its element, barely
      clinging to the hard, rock walls --
      much like the colony of bats
      which, once disturbed, begin to
      dive-bomb the intrepid
      speleologists. In this realm,
      geology rules.

      That is fine with Dean Smart, a
      cave consultant with the Royal
      Forestry Department (RFD) who
      is guiding the expedition. A
      geologist by training, Smart has
      been fascinated by caves ever
      since he was a boy.

      He seems at home in them,
      even enervated by them. At one
      point in Phra That Cave, he
      scrambles off through a thin,
      watery passage and is gone for
      10 minutes. But nobody seems
      alarmed. Elsewhere, fellow
      spelunker Bill Fanning climbs up
      through a narrow opening,
      before retreating in the face of
      foul air.

      Over the last three years, Smart
      has explored many impressive
      caves in Thailand, including
      Tham Sao Hin in Kanchanaburi,
      home to the tallest natural
      column in the world, around 61
      metres high.

      Now Smart intends to go a step
      further by carrying out a
      systematic study of hundreds of
      caves in Thung Yai Naresuan
      Wildlife Sanctuary, most of
      which have probably never
      been explored. The RFD project
      has already been promised
      financial support by the Thai
      Research Fund.

      ''Caves preserve things,''
      explains Smart, who hails from
      the Peaks District in England,
      where as a boy he used to
      explore many old lead mines.
      ''We've found teak coffins
      dating back 1,200 to 2,200
      years that have carvings from
      the beginning of Thailand's iron
      age. Out in the field, they would
      have been eaten by insects.

      ''So caves are important
      historically. Thai kings used to
      leave their signatures carved
      into cave walls, and some cave
      carvings in Ratchaburi prove
      that Dvaravati-style Buddhism
      was here before Theravada
      Buddhism.''

      Caves are also ecologically
      important, largely thanks to their
      constant environment.
      According to Smart, for
      instance, sediments preserved
      in caves contain a great deal of
      paleo-environmental information
      -- telling us about climatic
      conditions long ago -- probably
      even more than ice cores or
      deep-sea sediments which are
      usually studied by scientists.

      The steady nature of cave
      environments also means that
      the animals which become
      adapted to them are often
      unique and endemic. Smart
      cites the example of a
      one-millimetre long insect known
      as a springtail (from the genus
      troglopedetes ), which used to
      live all over Thailand. Then the
      climate changed, so that now,
      he says, you can find 12
      different species scattered in
      different caves in northern and
      central Thailand.

      By the same token, however,
      cave ecosystems are extremely
      fragile. In Ratchaburi, for
      instance, Tham Khao Bin -- a
      tourist site supervised by the
      Tourism Authority of Thailand
      and a local consortium of
      businessmen -- has been
      destroyed by bad management,
      according to Smart. Trails are
      poorly marked, and tourists
      wander where they like,
      trampling the fragile soil.

      ''They are very proud of their
      180,000-watt spotlights, but the
      lights are far too big and hot for
      the cave. As a result, the
      temperature inside is at least
      two degrees Celsius warmer
      than it should be,'' he maintains.
      ''This not only kills the animals --
      there are no more bats there,
      and a species of millipede which
      feed on their guano is gone, too
      -- but also increases the
      moisture in the air by sucking
      water out of the speleotherms
      [stalactites and stalagmites], so
      they start drying out.''

      One of the most interesting
      things about caves, it turns out,
      is observing how animals have
      adapted to the harsh
      environment. The most obvious
      adaptations involve the
      development of other senses
      besides eyesight to navigate in
      the pitch-black environment.

      Bats, of course, use sonar.
      They emit high-pitched sounds
      and then ''read'' the echoes to
      find their way around. So far,
      109 species of bats have been
      identified in Thailand -- including
      Kitti's Hog-nosed Bat, the
      world's smallest mammal -- and
      five of them were actually
      discovered here.

      Other species have developed
      their sense of touch. Some
      cave crickets have feelers 20
      times the length of their bodies,
      while ''Hairy Mary's'' ( scutigera
      centipedes) and cave-adapted
      catfish also sport impressive
      tendrils.

      Most species of fish have a
      lateral row of sensory to detect
      movements in the water around
      them, but those found in caves
      tend to have more of them.
      They are also usually blind and
      colourless. At least five species
      of cave fish are endemic to
      Thailand, says Smart, and two
      new species have been
      tentatively identified at a cave
      in Phitsanulok.

      ''Thailand has over 10 per cent
      of the world's known species of
      cave fish, and we hope to find
      some more,'' he says.

      In fact, that is the main reason
      for Borowsky's visit to Thailand.
      A biologist from New York
      University who specialises in
      genetics, Borowsky points out
      that caves are an excellent
      laboratory for studying both
      ecosystems and evolution.

      ''Cave ecosystems have far
      fewer variables than in the
      outside world, so it's easier to
      see how they interact,'' he
      explains. ''In terms of evolution,
      caves serve as a kind of
      repeated experiment to see
      what happens to animals who
      end up in an environment where
      there's no light and little food.''

      Researchers like Borowsky find
      that fish which have wandered
      into caves and bred there tend
      to undergo a process of
      convergent evolution: They lose
      their eyes and their skin colour,
      their sensory pits increase, and
      after a long time they lose their
      diurnal rhythm -- the body's
      natural sense of knowing when
      it is day and night -- and even
      their fear of predators, which
      are few and far between in cave
      streams.

      At first glance, it seems like a
      logical reaction. But think about
      it a minute. Although eyes may
      not be useful in caves, why
      should fish with fully formed
      eyes lose them over the
      generations? Put another way,
      what advantage do eyeless fish
      have over normal ones in
      caves?

      According to Borowsky, some
      theories suggest that, since it
      requires energy for a body to
      grow eyes, there is an
      advantage if that energy is put
      to some other use. Similarly,
      there is a theory that
      processing vision uses up
      valuable space in the brain,
      which again could be put to
      better use processing other
      senses.

      But Borowsky tends to support
      the hypothesis that complex
      systems naturally degenerate
      without the impetus of natural
      selection. In other words, all
      animals are randomly
      susceptible to bad mutations,
      and since cave fish don't need
      their eyes to survive, those fish
      born with poor or no eyesight
      don't get weeded out. The bad
      mutations accumulate through
      the generations, so that
      eventually cave fish can no
      longer see.

      ''This process is known as
      genetic drift, or random drift,
      and it's occurring all the time,''
      he says. ''Actually, genetic drift
      is what makes the process of
      DNA fingerprinting useful,
      because our random mutations
      mean that none of our genetic
      codes are identical [except in
      the case of identical twins].''

      Borowsky supports this
      ''entropy'' argument because it
      has been found that fish in
      caves near one another lose
      eyesight for completely different
      reasons. For instance, in some
      cases, the lens may have
      receded completely into the
      eyesocket, in others the nerves
      connecting the eye to the brain
      may no longer function.

      ''One species may lose the
      function of genes A, B and C.
      Another may lose D, E and F.
      But when the two types are
      hybridised, their offspring often
      have sight, because they have
      received all the necessary
      genes: A,B,C,D,E, and F,'' he
      explains.

      Borowsky has also carried out
      research in Mexico where he
      found something even more
      intriguing: cases where
      evolution has seemingly
      reversed itself. In one Mexican
      cave, a wall had collapsed
      letting sunlight in. Cave fish
      swimming under this karst
      window actually ''re-grew'' eyes.

      ''The case is interesting
      because there is a law, known
      as Dallo's Law, which states
      that the course of evolution
      cannot be reversed,'' notes
      Borowsky. This is because
      evolution is a random walk, so
      even if the environment reverts
      to its previous condition, it's
      highly unlikely that life will go
      back to ancient forms.

      ''But if Dallo's Law is violated in
      the case of Mexican cave fish,
      it would only be a
      misdemeanour, because the
      genetic vestiges of eyesight are
      still there, so it's easy for the
      fish to go back and regain their
      eyes,'' Borowsky concludes. 1